
26th Annual Meeting and Symposium of the
Desert Tortoise Council, March 16-18, 2001 Abstracts

Calcium and Phosphorus Availability in Native and
Exotic Food Plants
Lisa C. Hazard, Danielle R. Shemanski, Kenneth A. Nagy
Department of Organismic Biology, Ecology and Evolution, P.O. Box 951606, University of California, Los Angeles, CA 90095-1606

Exotic plants can comprise a major component of the diet for some
desert tortoises (Gopherus agassizii) in the Mojave Desert.
Introduced plants may not be as nutritious as native plants. Nutrient
availability in a native grass (Achnatherum hymenoides), an
introduced grass (Schismus barbatus), a native forb (Malacothrix
glabrata) and an introduced forb (Erodium cicutarium) were
measured in one to two year old juvenile desert tortoises. We fed
tortoises measured amounts of chopped foods daily for ~130 days (dry
grass) or ~90 days (fresh or thawed forb). Orts and feces were collected
daily and dried to constant mass, and calcium and phosphorus content of
food and feces were measured. Calcium digestibilities did not differ
significantly among diets. Because the grasses were lower in calcium
content, calcium availability (mg obtained/g dry food) was significantly
lower for grasses than forbs, and did not differ between grasses or
between forbs. Phosphorus digestibility was relatively high for the
forbs, but was negative for both grasses. Tortoises lost phosphorus
while feeding on the grasses, but gained phosphorus while eating forbs.
Calcium and phosphorus availability are better explained by the type of
food (forb vs. grass) than by its geographic origin (native vs. exotic);
however, if exotic grasses are replacing native forbs rather than native
grasses, the nutritional quality of the tortoises' diet could be
decreasing.
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