Survivorship was studied in an eastern Mojave Desert population
of desert tortoises from 1988 to 1996. The most reliable calculation
of maximum survival rates for tortoises >139 mm in carapace length,
based on survival of transmittered tortoises, was 0.934-1.00 in
most years . However, severe drought lowered survival of females
>186 mm in length to 0.696. Losses of 18.4-60 % of adult desert
tortoises during drought have also been observed in other rigorous
studies. Incorporating the longterm patterns of rainfall for
this site, I estimated that survivorship for females >188 mm was
0.907-0.964. Minimum nest depredation rates at this site ranged
from 18.4-42.0% of the annual egg production. The annualized
hatching rate based on three years study was 0.687, although sampling
artifacts suggest that the actual rate is 0.58. Incorporating
the most realistic population parameters of growth, survival by
life stage, and reproduction resulted in a lamda of 1.057 and
Ro of 4.35 for this population. Accurate estimates of survivorship
are critical for population behavior analyses. Survival rates
calculated from mark- recapture analyses, the lack of adequate
survival data for juvenile tortoises, and the lack of integration
of long-term variability in survivorship are discussed in the
context of the predictive ability of current population models
for the desert tortoise.