In 1996 and 1997, 102 tortoises were marked during two 45-day
censuses on a one square kilometer plot in the Tucson Mountains
as part of a three year project to study the desert tortoise at
Saguaro National Park. Frequency of observation of individual
tortoises was correlated with both body size and with average
distance from the edge of the plot. The relationship between body
size and number of prominent growth rings indicates a growth rate
similar to that obtained from recapture data if each ring is counted
as one year. Most tortoises greater than 20 years could not be
aged precisely because inner rings were worn away and outer rings
could not be resolved however it was clear that they were at least
20 years old. The age distribution was evaluated alternatively
by calculating a Lincoln Index estimate for each age class and
by using simple models to adjust the age frequency distribution
for effects of body size and distance to the plot boundary. Preliminary
results indicate possible high variability of annual productivity
and a low mortality rate for juvenile and immature tortoises.
The age distribution of tortoise shell remains recovered in the
Tucson Mountains indicates that egg and hatchling mortality is
highest although sample sizes are small. Validity of age determination
from growth rings needs to be tested in other populations where
many annual recaptures of young tortoises are expected. Methods
for the adjustment of age distributions need to be refined.